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“Organic Acids in Poultry Industry: A Review of Nutritional Advancements and Health Benefits.” World's Poultry Science Journal 80: 133–153

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Other authors suggested that gliadin was able to bind the mono-sialic ganglioside 1 (GM1) or the receptors for the proinflammatory chemokine 3 (CXCR3) in mouse models, leading to the release of zonulin which in turn decreases electrical resistance of epithelial layers, finally resulting in increased epithelial permeability (139, 255)